Evolution Reflected in Development

Below is an image of the developmental path from human conception to adult in comparison with evolutionary path from prokaryote to human.

Unlike Haeckel’s biogenetic law with its focus on physical forms, the comparison above also concerns activity, lifestyle and behaviour. Comparative stages may be vastly different in detail, but the similarity of general lifestyles and consecutive stages are there to be observed.

Human life begins in an aquatic environment. Toddlers gradually learn to walk upright from a previous state of crawling and moving around on all fours. The brains of children develop through daily interactions and experiences. This brain development accompanies the child’s increasing ability to achieve complex manipulation skills using hands that have been released from the task of providing support and locomotion, and also the practice of producing sounds using the various muscles of the mouth. Well developed brains allow for rational thinking and the creative use of language.

Human minds have brought about technological advances which have allowed human activities to engulf the planet. Signs of intelligent human activity are evident a good distance beyond the earth spreading ever further out into space.

The various forms of extant animals and all other life forms have evolved as an integral component of the living earth and the whole forms a dynamic system.

The various animal forms should be studied in the context of the complete system in both time and space.  Conditions would have been very different prior to the terrestrial colonization of earthly life In all probability none of the present aquatic animals would bear any resemblance to the aquatic ancestors of humans and other higher vertebrates save that at some stage they all require an aquatic environment for their continued existence.

From a point of view which regards physical organisms as the individual expressions of overarching general forms, the evolution of cetaceans need not have involved moving to the land only to return to the water at a later time. They may have reached the mammalian stage of evolution but in a way that was suitable for an aquatic lifestyle. They adopted the archetypal mammalian form in a way that suited an animal living in an aquatic environment and there would be no need to posit a terrestrial stage in their evolution.

It’s my belief that higher consciousness is ever present. Evolution is the process whereby higher forms of consciousness descend from the group level to the individual level. The most fully developed individual consciousness which I am aware of on earth can be found in humans but it is still rudimentary compared to the higher level group consciousness.

Plasticity is a fundamental feature of living systems at all levels from human brain development to the radiation of multicellular life. Paths are formed by branching out and becoming fixed along certain lines. It would be impossible to forecast specific paths but, nonetheless, there is a general overall direction.

Now that biological life has reached the stage where social organisms have become individually creative and rational, the all encompassing Word is reflected in single beings. This could not have come about without preparation and the evolution of earthly life is the evidence of this preparation. We, as individuals, are only able to use language and engage in rational thinking because our individual development has prepared us to do so. Likewise humanity could not arrive at the present state of culture without the evolutionary preparation in its entirety.

Focussing in at the lower level gives a picture of ruthless competition, of nature “red in tooth and claw”. But from a higher vantage point life benefits from this apparent brutality. For instance if a sparrowhawk makes regular hunting visits to a suitable habitat in your neighbourhood it signifies that this environment supports a healthy songbird population. In the case of the continued evolution of physical forms, survival of the breeding population is more important than any individual’s survival. In the evolution of consciousness the individual is the important unit.

I think it is a mistake to see biological evolution as a blind random groping towards an unknown and unknowable future.

896 thoughts on “Evolution Reflected in Development

  1. CharlieM: Bacteria are wise enough not to put all their eggs in one basket. The reason bacterial species are so successful is not because they all share a common genome, it is because they have a range of genomes with slight variation between them.

    Not the way I would have put it, but sure why not? Not sure why you restrict this to bacteria though. This applies to all living things. No variation, no evolution.

    CharlieM: We can see that there are different strains of coronavirus but they are all recognised as the same virus. Organisms can be individually different but of a common species or kind. Surely you can see this?

    CharlieM: I don’t need to name any because you have done it for me above.

    So here is the problem: HIV is a type of retrovirus, so do retroviruses make up a kind or is it a grouping of smaller kinds like HIV? Same goes for coronaviruses: Is it a kind in itself or a grouping of kinds like SARS-CoV-2?

    I think these are arbitrary groupings and that you will be unable to identify “kinds” united by “group wisdom” in there. We humans drew some convenient lines in the sand but nothing indicates viruses care for them. Why would they? Take the flu: There are multiple subtypes of influenza. Why would the “wisdom” of influenza A include influenza B and influenza C viruses but not those of infectious salmon anemia virus or Quaranjavirus? Because we humans named the former influenza and the latter not?

    CharlieM: Then some changes are preferentially generated. For example bacteria and archaea may use CRISPR-Cas systems to make changes within their own genomes.

    Yet mutations DNA sequence changes resulting from inaccurate replication by DNA polymerase are not preferentially generated, right? These can be reasonably described as errors, don’t you agree?

    CharlieM: We can leave out speculations about purpose, but we do know that the evolutionary process has brought about self consciousness and individuality.

    You have backpedaled so far that we are in agreement again. I do like to point out that the speculation is all yours.

  2. Allan Miller:
    CharlieM: Why can’t a reduction in fidelity also be a positive response for a colony?

    Allan Miller: It could. But if there is an active mechanism, I would expect it to be under some kind of specific control – some mechanistic relationship in which the stressor causes a consistent constitutional change which is not a byproduct, rather than the organism simply diverting resources and taking the mutagenic hit as collateral damage; an unavoidable secondary consequence. If a mutagenic chemical agent is introduced into a population, for example, is the population positively responding to it, or is its normal operation being interfered with? Now replace ‘mutagenic chemical agent’ with ‘stress’.

    Populations do not respond to external influences such as this. But individuals within the population react in different ways. It is this individual variability within a population which will determine if it can survive environmental change. But this fluctuation within populations bears little resemblance to what is seen as the great diversification of life over evolutionary time.

    CharlieM: Life depends on balance. Stress induces an imbalance. A loss of fidelity is also a gain in variability. This could be fatal for a high percentage of the individuals in a colony but it could also produce individuals which can tolerate the external disruption. Hence the colony can regrow and regain a balanced condition.

    Allan Miller: Mutations can be beneficial, you say? Gosh.

    The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other. But the fact that they add variety gives populations an adaptive advantage. Bacteria have remained as bacteria for millions of years by having this attribute.

    CharlieM: Alan Fox brought up the subject of equilibrium. Conditions within living systems must maintain a certain independence from their environment. We have selective barriers at all levels from our skin down to the barrel-like structures of some chaperone proteins.

    Allan Miller: I hope you’re not equivocating on the term ‘selective’. It has a very specific meaning in evolution, and it’s not that.

    I’m sure we all know that cell membranes are selective barriers in that they allow some, but not all molecules to pass through them by means of both passive and active transport.

    If I was referring to a specific type of selection I would have said so.

    Although now that you mention it I can see a resemblance to natural selection in which populations are regulated in a way that maintains their identity while the frequency of individual differences vary. 🙂

    CharlieM: At all these levels the systems must function in environments in which there are orderly, upbuilding forces and substances, also disruptive, destructive forces and substances. But just because some things are disruptive and destructive does not mean that cannot be beneficial.

    Allan Miller: You are describing something close to the ‘standard view’ of mutations.

    Good.

    CharlieM: Some living systems can tolerate high levels of environmental changes while others only narrow limits. Most Antarctic fish spend their lives within a very narrow environmental temperature range and will die of hyperthermia at temperatures above about 4 degrees Celsius.

    Allan Miller: You seem to be just waffling, here. It’s certainly true that organisms vary in the range of their environmental tolerances.

    The word niche is commonly used to describe the environment and habits of a species. In my opinion the niche of, say, a giant panda would be equivalent to a niche of an individual human rather than us as a species. To say that the human species has a niche is meaningless in my opinion.

    CharlieM: In peppered moths the ability to produce dark scales is very ancient. The typical form would not be peppered if areas of scales were not darkened. Even in the modern moths alteration to the cortex gene is only associated with a percentage of melanic forms. The estimated percentage is high but it is not total.

    Allan Miller: OK, and…?

    So why do I see all these headlines claiming that “the gene” for industrial melanism has been found?

  3. CharlieM: Populations do not respond to external influences such as this.

    I’d say they do – or rather, it’s a semantic nicety to say it’s not ‘the population’ that’s responding. An antibiotic kills a population by killing the individuals within it. Likewise, a mutagenic chemical agent, or a stress, changes the fidelity of replication of the individuals in the population, and hence the amount of variation in that population. Individuals don’t individually vary. Raising an “I think you’ll find…” finger when someone says that the mutagen changes the population is a bit pedantic. I realise I can talk about pedantry.

    But this fluctuation within populations bears little resemblance to what is seen as the great diversification of life over evolutionary time.

    The arc of individual development bears little resemblance to the great diversification of life over evolutionary time, but that doesn’t stop you beating that drum! I’m certainly not trying to draw any parallels between within-population change and between-population diversification. They are related, but not by analogy.

    The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other. But the fact that they add variety gives populations an adaptive advantage. Bacteria have remained as bacteria for millions of years by having this attribute.

    Mutations are a part of stasis, now? Interesting …

    If I was referring to a specific type of selection I would have said so.

    Possibly. I was just checking.

    Although now that you mention it I can see a resemblance to natural selection in which populations are regulated in a way that maintains their identity while the frequency of individual differences vary.

    Interesting that this is what you think natural selection does.

    The word niche is commonly used to describe the environment and habits of a species. In my opinion the niche of, say, a giant panda would be equivalent to a niche of an individual human rather than us as a species. To say that the human species has a niche is meaningless in my opinion.

    The word ‘niche’ suffers from similar problems to the word ‘gene’. It can be somewhat labile. Nonetheless, I have never seen anyone use it to refer to individuals.

    So why do I see all these headlines claiming that “the gene” for industrial melanism has been found?

    No idea. You must be more widely read than I; this has escaped my notice.

  4. CharlieM,

    The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other.

    Ooh! Ooh! I let that one slide! This is flat-out wrong in its second part. The inheritance of mutation in a finite world prevents any population experiencing it from ‘standing still’. All classes of mutation beyond the substantially detrimental have the capacity to have a lasting effect: for the population at time t2 to differ irreversibly from that at t1. It’s something you can’t turn off.

  5. Corneel:
    Me (Corneel): There is no such thing as naked DNA, or so I’ve been told. It is the whole of the cell that is expressing the selfish DNA bits, no?

    Charlie: I do not believe that selfishness applies to nucleotide patterns.

    Corneel: I didn’t say that selfishness applies to nucleotide patterns (though it does, metaphorically speaking). I said that if you were being consistent, you’d have accepted that it is the whole of the cell that is displaying selfish behaviour, not just the DNA.

    Should I believe our hands are being selfish because their form is copied in our descendants?

    Me (Corneel): It appears you forgot that you specified a scenario where organisms couldn’t die.

    Charlie: Only in order to demonstrate the absurdity of the consequences.

    Corneel: No, you introduced this scenario to argue that the death of organisms was a requirement for evolution to occur, which is false. You also suggested that the death of organisms is in some way analogous to apoptosis, which is also false.

    Can you explain how natural selection could occur without death?

    The death of mature mayflies leaves room for future populations. The apoptosis of cells during vertebrate limb development leave room for digits to form. Two levels in which the death of individuals benefits the higher entity to which they belong. (Cell death in one case and the death of individual organisms in the other.)

    CharlieM: For multicellular creatures like us germline cells have the capability to be heritable and so instigating lines of descent of individual organisms (evolution). Somatic cells have the capacity to create lines of descent of cells within a developing individual organism (development).

    Corneel: Close enough. So consider this: If somatic cells lost their ability to execute apoptosis and if this change would be mitotically heritable, what would be the chance this change would end up in the offspring? And what would happen to the chances the poor organism these cells were part of would leave any offspring?

    Apoptosis is a natural process within individual development. I don’t believe any embryo would reach full term without it, let alone reaching sexual maturity.

    CharlieM: That example was in response to your implication that continued existence and a sense of self were mutually dependent on each other. Why else would you ask. What does continued existence mean to something that does not possess a sense of self?

    Corneel: And you replied that meaning came from the observer, not from buildings or cells. So when I asked: “So if apoptosis is deleterious for the cell, then what exactly is being sacrificed?”, the answer should have been: “Absolutely nothing is sacrificed from the perspective of the cell, but Charlie felt bad for it anyway.”

    Guess what? Corneel doesn’t pity this cell at all, ’cause apoptosis is not deleterious. The word “deleterious” has a specific meaning in genetics: it means lowering fitness or predisposing to disease. Apoptosis doesn’t do either of those things. Hence, apoptosis is not deleterious to a cell.

    If the proteins within a cell are indiscriminately degraded how can you say that this does not lower the fitness of the cell?

    CharlieM: Then why didn’t you say it’s only a select set of viruses that you don’t want me to bother harbouring?

    Cornell: I am pretty sure I mentioned infection. The viruses that infect you are definitely not going to give you good times. It is you that views all viruses as a big happy family helping each other for the benefit of all viruskind.

    I’d like to clarify that your view of my view is a false view of my view and so it is not my view which views viruses as being very diverse. See? 🙂

    There is no mutual cooperation among viruses that I know of.

  6. CharlieM: Should I believe our hands are being selfish because their form is copied in our descendants?

    Silly billy. Corneel wrote

    It is the whole of the cell that is expressing the selfish DNA bits, no?

    Thus ‘you should believe’ that your hands are merely tools that your DNA is using to express their (the DNA’s) selfishness. There’s a Dawkins bloke who wrote about this way of thinking a while back.

    CharlieM: Can you explain how natural selection could occur without death?

    Differential reproduction. This, too, has been covered before.

    CharlieM: If the proteins within a cell are indiscriminately degraded how can you say that this does not lower the fitness of the cell?

    ‘Fitness’ is a technical term that does not apply to somatic cells, that’s how.

    CharlieM: There is no mutual cooperation among viruses that I know of.

    I am guessing that you haven’t looked very hard.
    E4 kinder link

  7. Corneel:
    CharlieM: Bacteria are wise enough not to put all their eggs in one basket. The reason bacterial species are so successful is not because they all share a common genome, it is because they have a range of genomes with slight variation between them.

    Corneel: Not the way I would have put it, but sure why not? Not sure why you restrict this to bacteria though. This applies to all living things. No variation, no evolution.

    Giving specific examples does not mean I’m being restrictive. I’m just being selective in what I am focusing on. 🙂

    CharlieM: We can see that there are different strains of coronavirus but they are all recognised as the same virus. Organisms can be individually different but of a common species or kind. Surely you can see this?

    CharlieM: I don’t need to name any because you have done it for me above.

    Corneel: So here is the problem: HIV is a type of retrovirus, so do retroviruses make up a kind or is it a grouping of smaller kinds like HIV? Same goes for coronaviruses: Is it a kind in itself or a grouping of kinds like SARS-CoV-2?

    I’m sure you’ve heard of nested hierarchies. As is usual in living systems, viruses (too) have been classed taxonomically.

    Corneel: I think these are arbitrary groupings and that you will be unable to identify “kinds” united by “group wisdom” in there. We humans drew some convenient lines in the sand but nothing indicates viruses care for them. Why would they? Take the flu: There are multiple subtypes of influenza. Why would the “wisdom” of influenza A include influenza B and influenza C viruses but not those of infectious salmon anemia virus or Quaranjavirus? Because we humans named the former influenza and the latter not?

    I don’t know enough about the relationships between viruses to judge. I use the word “kinds” to describe groups of which individuals are members of. It is a broader category which may or may not align with a level of classification such as species or genera.

    For example the species influenza A would include all influenza A individuals.

    CharlieM: Then some changes are preferentially generated. For example bacteria and archaea may use CRISPR-Cas systems to make changes within their own genomes.

    Corneel: Yet mutations DNA sequence changes resulting from inaccurate replication by DNA polymerase are not preferentially generated, right? These can be reasonably described as errors, don’t you agree?

    Yes they could be seen as copying errors.

    CharlieM: We can leave out speculations about purpose, but we do know that the evolutionary process has brought about self consciousness and individuality.

    You have backpedaled so far that we are in agreement again. I do like to point out that the speculation is all yours.

    To say that evolution has produced individual self-consciousness is not speculation, it is an observation.

    To say that this has happened by chance or to say that it was destined to be are equally speculative for most of us.

  8. Jock already fielded some stuff, I see. Starting on the remaining issues:

    CharlieM: mayflies

    Yeah, that made me smile. Though the life span of mayflies may be ephemeral, their death is not programmed. Most species of mayflies do not have functional mouthparts or digestive system, so they simply waste away.

    Nice try though.

    CharlieM: Apoptosis is a natural process within individual development. I don’t believe any embryo would reach full term without it, let alone reaching sexual maturity.

    Exactly, so somatic cells that appropriately execute apoptosis prevent the termination of the lineage they are part of by way of their close relationship to germline cells. That’s not exactly deleterious, is it?

    CharlieM: I’d like to clarify that your view of my view is a false view of my view and so it is not my view which views viruses as being very diverse.

    ahem:

    If a virus can change the DNA of its host in order to ensure the continued existence of its kind, would that not be a wise move on the part of the virus? The fact that viruses mutate indicates that it is not the DNA sequence but the virus species or kind that is being preserved. Their DNA varies which enables them to survive as a group.

    If viruses do not give a rat’s arse about each other, then why would it be “wise” to “ensure the continued existence of its kind”? Looks like a big happy virus family to me.

  9. CharlieM: I’m sure you’ve heard of nested hierarchies. As is usual in living systems, viruses (too) have been classed taxonomically.

    I do not see how “group wisdom” stacks hierarchically.

    CharlieM: I don’t know enough about the relationships between viruses to judge. I use the word “kinds” to describe groups of which individuals are members of.

    The problem is the concept of “kind” in the mind of the average layperson. It has acquired a lot of baggage, you see?

    CharlieM: Yes they could be seen as copying errors.

    FINALLY!

    CharlieM: To say that evolution has produced individual self-consciousness is not speculation, it is an observation.

    To say that this has happened by chance or to say that it was destined to be are equally speculative for most of us.

    I like this quote by KN. I believe it has some relevance here as well:

    I like to frame the issue as follows: there is no evidence for an empirically detectable mechanism that first determines what traits would be advantageous and then brings those traits about. So in that sense evolution is “unguided”.

    Whether evolution is “unguided” in any more metaphysically demanding sense, in a sense that would have any relevance for theism or atheism, is not a question that can be addressed in scientific terms.

  10. Allan Miller:
    CharlieM: Populations do not respond to external influences such as this.

    Allan Miller: I’d say they do – or rather, it’s a semantic nicety to say it’s not ‘the population’ that’s responding. An antibiotic kills a population by killing the individuals within it. Likewise, a mutagenic chemical agent, or a stress, changes the fidelity of replication of the individuals in the population, and hence the amount of variation in that population. Individuals don’t individually vary. Raising an “I think you’ll find…” finger when someone says that the mutagen changes the population is a bit pedantic. I realise I can talk about pedantry.

    For me evolution is a well balanced journey. Natural selection maintains the balance but it does not control the route. For the living world to persist there must be a balance between life and death. And persistence of life at higher levels must include death at lower levels. A multicellular organism or a colony of single celled organisms maintains its life through death at the cellular level. A higher taxonomic group maintains it life through death of the individual organisms within it. A kingdom of nature maintains its life through the death of taxonomic groups within it. And as far as I’m aware earthly life as a whole has yet to reach the stage of total death even if death is a constant feature of lower levels.

    So, arguing against myself, I could say that in one sense I am a population of cells responding to external stimuli.

    But this fluctuation within populations bears little resemblance to what is seen as the great diversification of life over evolutionary time.

    Allan Miller: The arc of individual development bears little resemblance to the great diversification of life over evolutionary time, but that doesn’t stop you beating that drum! I’m certainly not trying to draw any parallels between within-population change and between-population diversification. They are related, but not by analogy.

    They are related fractally as in the whole reflected in the parts.

    CharlieM: The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other. But the fact that they add variety gives populations an adaptive advantage. Bacteria have remained as bacteria for millions of years by having this attribute.

    Allan Miller: Mutations are a part of stasis, now? Interesting …

    Nothing is truly static. Analogy alert! Jupiter’s great red spot is a very dynamic system but it has been consistently present for at least hundreds of years. A fairly stable feature which at lower levels is a very energetic atmospheric storm.

    If I was referring to a specific type of selection I would have said so.

    Allan Miller: Possibly. I was just checking.

    Phoodoo will be pleased. 🙂

    CharlieM: Although now that you mention it I can see a resemblance to natural selection in which populations are regulated in a way that maintains their identity while the frequency of individual differences vary.

    Allan Miller: Interesting that this is what you think natural selection does.

    Typical and melanic peppered moths keep their identity as peppered moths while natural selection causes fluctuations in their relative densities. Similar processes can be seen in Galapagos finches. These examples are a favourite argument for natural selection in action.

    CharlieM: The word niche is commonly used to describe the environment and habits of a species. In my opinion the niche of, say, a giant panda would be equivalent to a niche of an individual human rather than us as a species. To say that the human species has a niche is meaningless in my opinion.

    Allan Miller: The word ‘niche’ suffers from similar problems to the word ‘gene’. It can be somewhat labile. Nonetheless, I have never seen anyone use it to refer to individuals.

    Maybe not in the case of bacteria, but have you never heard it being said that someone has found their niche?

    CharlieM: So why do I see all these headlines claiming that “the gene” for industrial melanism has been found?

    Allan Miller: No idea. You must be more widely read than I; this has escaped my notice.

    I didn’t find this by being widely read, I found it through focused searches on industrial melanism.

  11. Corneel:CharlieM: Bacteria have remained as bacteria for millions of years by having this attribute.

    Corneel: No, they met their demise.

    What? The eubacteria kingdom is no more? It is a dead kingdom? It is pushing up the daisies? Where do I send the flower to? 🙁

  12. CharlieM:

    They are related fractally as in the whole reflected in the parts.

    Nope, not that either. They are related because change in lineage (anagenesis) is the direct cause of divergence between two (cladogenesis), given a reproductive barrier.

    Nothing is truly static. Analogy alert! Jupiter’s great red spot is a very dynamic system but it has been consistently present for at least hundreds of years. A fairly stable feature which at lower levels is a very energetic atmospheric storm.

    Questionable relevance alert.

    Phoodoo will be pleased.

    I’ve rarely known phoodoo be pleased by anything.

    Maybe not in the case of bacteria, but have you never heard it being said that someone has found their niche?

    Ah, good old semantics, our constant companion. This is not the ecological use of the term.

    …focused searches on industrial melanism

    I’ve just done a focused search and found that the difference is attributed to a transposable element. Is this reasonably called the gene ‘for’ melanism? Is that how the authors present it, or journalists? I’d be interested in a well-defended rationale either way.

  13. CharlieM:
    Typical and melanic peppered moths keep their identity as peppered moths while natural selection causes fluctuations in their relative densities. Similar processes can be seen in Galapagos finches. These examples are a favourite argument for natural selection in action.

    Natural selection in action at any point is progressing towards the elimination of one form and fixation of the other, as a logical endpoint of the current trend. Of course that trend could change, leading to ‘fluctuation’, but you err in thinking that fluctuation is a component of selection.

  14. CharlieM: What? The eubacteria kingdom is no more? It is a dead kingdom? It is pushing up the daisies? Where do I send the flower to? 🙁

    I am given to understand that one does not send flowers, but potatoes.

  15. Allan Miller:
    CharlieM: The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other

    Allan Miller: Ooh! Ooh! I let that one slide! This is flat-out wrong in its second part. The inheritance of mutation in a finite world prevents any population experiencing it from ‘standing still’. All classes of mutation beyond the substantially detrimental have the capacity to have a lasting effect: for the population at time t2 to differ irreversibly from that at t1. It’s something you can’t turn off.

    I’ll happily go along with that. I went a bit far saying the noise has no effect.

    Michael Behe argues that the vast majority of mutations that do have adaptive effects and alter organisms through Darwinian processes, do so by degrading genes that already exist. In this video he cites examples such as polar bears and dogs.

    Considering your point that mutations can have irreversible effects. In the case of peppered moths, just think if the supremacy of the melanic form had gone as far as to annihilate the other forms. Through Darwinian processes at time t1 the species would have consisted of a wide range of patterns among individuals compared to time t2 at which point individuals would be restricted to the darkened form.

    This is not a good argument for the power of Darwinian evolution.

  16. Corneel:
    CharlieM: I’m sure you’ve heard of nested hierarchies. As is usual in living systems, viruses (too) have been classed taxonomically.

    Corneel: I do not see how “group wisdom” stacks hierarchically.

    I do see how there can be an orderly progression. And the lowest, most nascent form of wisdom is individual conscious human wisdom.

    CharlieM: I don’t know enough about the relationships between viruses to judge. I use the word “kinds” to describe groups of which individuals are members of.

    Corneel: The problem is the concept of “kind” in the mind of the average layperson. It has acquired a lot of baggage, you see?

    Yes I can see problems when people attach their own prejudices to the use of a word.

    CharlieM: Yes they could be seen as copying errors.

    Corneel: FINALLY!

    And so there is a polarity between the disruption of copying errors and error correction processes which tries to maintain order.

    The wisdom lies not in eliminating errors completely but in using them constructively.

    CharlieM: To say that evolution has produced individual self-consciousness is not speculation, it is an observation.

    To say that this has happened by chance or to say that it was destined to be are equally speculative for most of us.

    Corneel: I like this quote by KN. I believe it has some relevance here as well:

    I like to frame the issue as follows: there is no evidence for an empirically detectable mechanism that first determines what traits would be advantageous and then brings those traits about. So in that sense evolution is “unguided”.

    Whether evolution is “unguided” in any more metaphysically demanding sense, in a sense that would have any relevance for theism or atheism, is not a question that can be addressed in scientific terms.

    What does it mean for a trait to be advantageous? If by advantageous it means that they allow organisms the most reproductive success then bacteria and viruses by far outcompete us mere humans. There are no higher animals on the planet that can match bacteria and viruses if that is the measure of successful evolution.

    Goethean science is not an activity which speculates about evolutionary origins. It is more concerned with observing the natural world and its processes as it is experienced.

    Goethe said:

    “Light has called forth one organ to become its like, and thus the eye is formed by the light and for the light so that the inner light may emerge to meet the outer light.”

    If we look at the relationship between light and life, light plays a vital role in the growth of organic compounds. In the case of plants, light enables them to grow living substance from inorganic compounds. And this in turn leads to an advancement of the relationship between light and life.

    As Goethe said:

    “…the eye has to thank the light for its existence. The light calls forth out of indifferent auxiliary animal organs, an organ that is akin to itself; the eye forms itself by the light for the light, so that the inner light can meet the external light.”

    It is obvious that without light eyes would not have evolved. And sense organs lead to, not only an awareness of the external world, but an awareness of self in relation to the world.

    There is a progression in evolution which matches the progression we observe in the development of the individual human. From growth/expansion through the birth of sentience on to the arrival of self-awareness.

    Self-awareness is a feature of life which does not give us outstandingly better reproductive rates over relatively more simple prokaryotes, yet here we are. We have been given access to an inner light that can say, “I see with my understanding”. Was this just an accident of blind evolution?

  17. CharlieM: What does it mean for a trait to be advantageous? If by advantageous it means that they allow organisms the most reproductive success then bacteria and viruses by far outcompete us mere humans.

    True, but let’s be honest: you haven’t been trying very hard, have you?

    CharlieM: There is a progression in evolution which matches the progression we observe in the development of the individual human. From growth/expansion through the birth of sentience on to the arrival of self-awareness.

    Whatever floats your boat!

    CharlieM: Self-awareness is a feature of life which does not give us outstandingly better reproductive rates over relatively more simple prokaryotes, yet here we are. We have been given access to an inner light that can say, “I see with my understanding”. Was this just an accident of blind evolution?

    That’s my take on it. As I said: you may disagree but there is no empirical evidence that evolution is steering to some specific goal.

  18. CharlieM,

    Michael Behe …

    Can go take a running jump. Who he? Where’s his rigorous analysis of relative adaptiveness been published?

    Considering your point that mutations can have irreversible effects. In the case of peppered moths, just think if the supremacy of the melanic form had gone as far as to annihilate the other forms. Through Darwinian processes at time t1 the species would have consisted of a wide range of patterns among individuals compared to time t2 at which point individuals would be restricted to the darkened form.

    This is not a good argument for the power of Darwinian evolution.

    If the melanic form does better than all rivals under the environmental conditions prevalent at the time, and ends up being the sole form at t2 as a consequence, I’m not seeing the problem. Melanic is best; lo! melanic it is! Good old Natural Selection.

    ‘Darwinian’ selection requires variation, but is not defeated by the observation that variation diminishes as selection progresses. It’s kind of expected.

  19. Allan Miller:
    CharlieM: They are related fractally as in the whole reflected in the parts.

    Allan Miller: Nope, not that either. They are related because change in lineage (anagenesis) is the direct cause of divergence between two (cladogenesis), given a reproductive barrier.

    Not surprisingly we see things from different angles.

    CharlieM: Nothing is truly static. Analogy alert! Jupiter’s great red spot is a very dynamic system but it has been consistently present for at least hundreds of years. A fairly stable feature which at lower levels is a very energetic atmospheric storm.

    Allan Miller Questionable relevance alert.

    What holds for one level of observed reality does not necessarily hold at all levels.

    CharlieM: Phoodoo will be pleased.

    Allan Miller: I’ve rarely known phoodoo be pleased by anything.

    Do you know phoodoo well enough to judge?

    CharlieM: Maybe not in the case of bacteria, but have you never heard it being said that someone has found their niche?

    Allan Miller: Ah, good old semantics, our constant companion. This is not the ecological use of the term.

    Surely ecologists have borrowed this term from general use for a reason?

    CharlieM…focused searches on industrial melanism

    Allan Miller: I’ve just done a focused search and found that the difference is attributed to a transposable element. Is this reasonably called the gene ‘for’ melanism? Is that how the authors present it, or journalists? I’d be interested in a well-defended rationale either way.

    Some, but not all, of the differences are associated in some way with a specific gene. Is that gene specific to melanism?

  20. CharlieM: Not surprisingly we see things from different angles.

    But that is how they are related! Can you envisage uncorrelated change in 2 separate populations leading to anything other than divergence?

    What holds for one level of observed reality does not necessarily hold at all levels.

    Rendering your too-frequent resort to analogy somewhat undermined!

    Do you know phoodoo well enough to judge?

    I don’t need to in order to make the statement “I’ve rarely known phoodoo be pleased by anything”. It was, after all, a joke.

    Surely ecologists have borrowed this term from general use for a reason?

    Of course – in order to permit endless equivocation! Scientists are always faced with 2 choices – generate an impenetrable neologism, or add a meaning to an existing word. The fun we have when they do the latter!

    Some, but not all, of the differences are associated in some way with a specific gene. Is that gene specific to melanism?

    Don’t know. Would it be the gene ‘for’ melanism if it was?

  21. Allan Miller:
    CharlieM: Typical and melanic peppered moths keep their identity as peppered moths while natural selection causes fluctuations in their relative densities. Similar processes can be seen in Galapagos finches. These examples are a favourite argument for natural selection in action.

    Allan Miller: Natural selection in action at any point is progressing towards the elimination of one form and fixation of the other, as a logical endpoint of the current trend. Of course that trend could change, leading to ‘fluctuation’, but you err in thinking that fluctuation is a component of selection

    Were the melanic forms not fixed in the population before selection due to tree discolouration caused by industrial pollution? Fluctuation in relative numbers of these previously fixed traits comes about due to selective pressures does it not?

    Say natural selection brought about the elimination of all peppered moths except the melanic forms. Surely this is a restriction of variability which will tend to make the species less capable of adapting to future changing conditions?

  22. Corneel:
    CharlieM: What? The eubacteria kingdom is no more? It is a dead kingdom? It is pushing up the daisies? Where do I send the flower to?

    Corneel: I am given to understand that one does not send flowers, but potatoes.

    You don’t happen to have the phone number for “Interpatata” do you? 🙂

  23. CharlieM: You don’t happen to have the phone number for “Interpatata” do you?

    You are lucky that I got that joke; that company goes by a very different name here.

    Anyways, the point is that the bacteria from millions of years ago met their demise sensu Charlie, just like theropod dinosaurs did. Modern bacteria are as different from their mesozoic cousins as modern birds are from mesozoic theropod dinosaurs, contra your claim that “The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other.”.

    Either that, or concede that eukaryotes have remained as eukaryotes for millions of years. Now you are just engaging in another round of “all those bacteria look the same to me”.

  24. Corneel:
    CharlieM: What does it mean for a trait to be advantageous? If by advantageous it means that they allow organisms the most reproductive success then bacteria and viruses by far outcompete us mere humans.

    Corneel: True, but let’s be honest: you haven’t been trying very hard, have you?

    I’m no bacterium but I have had my reproductive successes. 🙂

    CharlieM: There is a progression in evolution which matches the progression we observe in the development of the individual human. From growth/expansion through the birth of sentience on to the arrival of self-awareness.

    Corneel: Whatever floats your boat!

    Displacement floats my boat.

    CharlieM: Self-awareness is a feature of life which does not give us outstandingly better reproductive rates over relatively more simple prokaryotes, yet here we are. We have been given access to an inner light that can say, “I see with my understanding”. Was this just an accident of blind evolution?

    Corneel: That’s my take on it. As I said: you may disagree but there is no empirical evidence that evolution is steering to some specific goal.

    Below is the text which accompanies the attached figure.

    The tree of life—an illustration of the major branches of life. A geological timescale moves radially from the bottom to the top of the diagram. All life on Earth is related. To better comprehend this reality, some of the major organismal groups are illustrated with colored branches for simplicity. The underlying layer of gray branches implies a more realistic and chaotic interconnectedness of life’s lineage. The letters a–g denote the locations of common ancestors, including those of plants (b) and of multicellular organisms (a). Many of the common ancestors of aceols and flatworms, insects, vertebrates, and land animals (annelids, arthropods, mollusks, echinoderms, and vertebrates) (c–f) can be traced to the Cambrian explosion of diversity.

    According to that tree bacteria have only themselves as common ancestors whereas humans have an (a) to (g) of common ancestors. If evolution is defined as change over time then humans have evolved further than bacteria.

    Even if evolution isn’t seen as progressing towards a specific goal it can been that up to this time there has been a progression towards creatures which possess a higher individual consciousness.

  25. CharlieM,

    Here is a tree of life that better represents modern understanding of relative diversity within Bacteria, Archaea and Eukarya. For giggles, try to locate “ancestors (a) to (g)” from your figure. Hopefully, you’ll realize that a fair amount of navelgazing went into its production.

    ETA: Also note how, in your figure, there is an implied progression from left to right. Phylogenetic trees can be rotated around their internal nodes without changing the topology, so why was it represented this way? Can you guess?

  26. CharlieM: Were the melanic forms not fixed in the population before selection due to tree discolouration caused by industrial pollution? Fluctuation in relative numbers of these previously fixed traits comes about due to selective pressures does it not?

    The environmental change caused a change in the direction of selection. This does not mean that fluctuation is a component of selection, simply that selection is responsive to it.

    Say natural selection brought about the elimination of all peppered moths except the melanic forms. Surely this is a restriction of variability which will tend to make the species less capable of adapting to future changing conditions?

    Sure. It happens. But it isn’t a universal. There is a constant flux of new mutations at one end, and a constant flux of fixation due to drift and selection at the other. So no biggie.

  27. Allan Miller:
    CharlieM, Michael Behe …

    Allan Miller: Can go take a running jump. Who he?

    That’s one way of keeping fit. I’m using “fit” in its general, not necessarily evolutionary, sense. 🙂

    Allan Miller: Where’s his rigorous analysis of relative adaptiveness been published?

    I don’t know. But even if he had carried out and published his own experiments on this, it would not give enough data. In proposing that the vast majority of beneficial mutations are degrative he has examined and reviewed the literature to gather information from a much wider source than he alone could have produced.

    CharlieM: Considering your point that mutations can have irreversible effects. In the case of peppered moths, just think if the supremacy of the melanic form had gone as far as to annihilate the other forms. Through Darwinian processes at time t1 the species would have consisted of a wide range of patterns among individuals compared to time t2 at which point individuals would be restricted to the darkened form.

    This is not a good argument for the power of Darwinian evolution.

    Allan Miller: If the melanic form does better than all rivals under the environmental conditions prevalent at the time, and ends up being the sole form at t2 as a consequence, I’m not seeing the problem. Melanic is best; lo! melanic it is! Good old Natural Selection.

    Melanic might be best for the individual, but narrows the options available to the group. Having the option of various forms to cope with changing environments is best for the population.

    Allan Miller: ‘Darwinian’ selection requires variation, but is not defeated by the observation that variation diminishes as selection progresses. It’s kind of expected.

    I am not trying to defeat Darwinian selection, I am just pointing out its limitations.

  28. CharlieM:
    I don’t know. But even if he had carried out and published his own experiments on this, it would not give enough data. In proposing that the vast majority of beneficial mutations are degrative he has examined and reviewed the literature to gather information from a much wider source than he alone could have produced.

    And what protection do we mere mortals have from any suspicion he might have cherry-picked? He is, after all, an IDist. I realise that’s a reverse argument from authority, but that’s all you’re doing.

    Melanic might be best for the individual, but narrows the options available to the group. Having the option of various forms to cope with changing environments is best for the population.

    Isn’t it fortunate that variation is, indeed, the rule? Except where it isn’t. We get along just fine with everyone having a nose, for example.

    But here’s the thing: what would you call the response to changing environments, given population variation with the capacity to so respond? Huh? It’s got a name, you know…

    I am not trying to defeat Darwinian selection, I am just pointing out its limitations.

    Thank goodness you came! Otherwise people might have run away with the idea that Natural Selection was all there was to evolution.

  29. Allan Miller: Isn’t it fortunate that variation is, indeed, the rule? Except where it isn’t. We get along just fine with everyone having a nose, for example.

    If the entire group does it, it’s group wisdom. Except where some are doing something different which is group wisdom as well.

    If they wouldn’t be doing it, they wouldn’t be doing it. Things’d be different then and we can clearly observe they are not. Things are definitely like they are now.

  30. Corneel,

    There is also a creative circularity in deprecating Natural Selection for its reduction of the variation whose principal purpose is to provide a response to … Natural Selection.

  31. Allan Miller: There is also a creative circularity in deprecating Natural Selection for its reduction of the variation whose principal purpose is to provide a response to … Natural Selection.

    Yeah like: “You shouldn’t drive your car because then all the fuel disappears”.

  32. Allan Miller:
    CharlieM: Not surprisingly we see things from different angles.

    Allan Miller: But that is how they are related! Can you envisage uncorrelated change in 2 separate populations leading to anything other than divergence?

    In the case of the peppered moth we can imagine two extreme forms diverging and remaining separate. But where will this lead? It is believed that moths existed on earth around 200 million years ago, so it is a fair assumption that these diverging peppered moths are descendants of lines of moths stretching back at least 200 million years. There is no evidence that, should they continue to exist, they will be anything but moths in the future. A creature whose form has been recognizable as a moth for such a long time has obviously made quite a specialistic, stable, evolutionary path for itself.

    CharlieM: What holds for one level of observed reality does not necessarily hold at all levels.

    Allan Miller: Rendering your too-frequent resort to analogy somewhat undermined!

    Not at all.

    If we recognize what Goethe was saying about everything changing yet remaining the same, we can see essential unity observed diversity. The use of light made by the leaf of a plant cannot be likewise attributed to the petals of the flower, but Goethe points to the essential unity of leaf and petal.

    From I.E.P.

    Goethe’s morphology, in opposition to the static taxonomy of Linnaeus, studied these perceptible limitations not merely in order to classify plants in a tidy fashion, but as instances of natural generation for the sake of intuiting the inner working of nature itself, whole and entire. Since all organisms undergo a common succession of internal forms, we can intuitively uncover within these changes an imminent ideal of development, which Goethe names the ‘originary phenomenon’ or Urphänomen. These pure exemplars of the object in question are not some abstracted Platonic Idea of the timeless and unchanging essence of the thing, but “the final precipitate of all experiences and experiments, from which it can ever be isolated. Rather it reveals itself in a constant succession of manifestations,” (Goethe 1981, 13: 25). The Urphänomen thus offer a sort of “guiding thread through the labyrinth of diverse living forms,” (Goethe 1961-3, 17: 58), which thereby reveals the true unity of the forms of nature in contrast to the artificially static and lifeless images of Linneaus’ system. Through the careful study of natural objects in terms of their development, and in fact only in virtue of it, we are able to intuit morphologically the underlying pattern of what the organic object is and must become. “When, having something before me that has grown, I inquire after its genesis and measure the process as far back as I can, I become aware of a series of stages, which, though I cannot actually see them in succession, I can present to myself in memory as a kind of ideal whole,”

    Organs at different levels of development might serve very different purposes but still retain an essential unity.

    CharlieM: Do you know phoodoo well enough to judge?

    Allan Miller: I don’t need to in order to make the statement “I’ve rarely known phoodoo be pleased by anything”. It was, after all, a joke.

    And I was just making an off the cuff remark.

    CharlieM: Surely ecologists have borrowed this term from general use for a reason?

    Allan Miller: Of course – in order to permit endless equivocation! Scientists are always faced with 2 choices – generate an impenetrable neologism, or add a meaning to an existing word. The fun we have when they do the latter!

    I think the concept of “niche” has been used to try to quantify relationships within ecosystems. But niches can have so many dimensions that to talk about an organisms niche in general becomes very abstract. Occupation of niches can be viewed with regard to temperature, salinity, relationships, feeding habits, and a host of other factors. Unlike humans, most species have a fairly common set of niche parameters between individuals. Niches for humans is often a matter of individual choice.

    CharlieM: Some, but not all, of the differences are associated in some way with a specific gene. Is that gene specific to melanism?

    Allan Miller: Don’t know.

    From what I can remember that gene is not specific to melanism, but I don’t have time to look just now.

    Allan Miller: Would it be the gene ‘for’ melanism if it was?

    No, even then I would not call it the gene for melanism.

  33. CharlieM: In the case of the peppered moth we can imagine two extreme forms diverging and remaining separate. But where will this lead?

    Speciation is the word you’re looking for.

  34. Corneel: Anyways, the point is that the bacteria from millions of years ago met their demise sensu Charlie, just like theropod dinosaurs did. Modern bacteria are as different from their mesozoic cousins as modern birds are from mesozoic theropod dinosaurs, contra your claim that “The vast majority of mutations are just incidental “noise” which does not have any lasting affect one way or the other.”.

    Either that, or concede that eukaryotes have remained as eukaryotes for millions of years. Now you are just engaging in another round of “all those bacteria look the same to me”.

    As my previous post demonstrates I’ve never disputed that eukaryotes have remained eukaryotes for millions of years.

    To get a clue as to how different organisms have changed over their evolution we can look at individual development. Bacteria which reproduce by cell division are born mature with all the attributes of the parent cell. Now consider bird reproduction. The offspring begins life as a fertilized egg and from then on it matures adding differentiated cells, various organs, tissues and structures as it develops.

    You might believe that individual bacteria have their own characteristics and personalities but it’s not a belief that I share.

  35. CharlieM: You might believe that individual bacteria have their own characteristics and personalities but it’s not a belief that I share.

    So bacteria do have inter-individual variation that allows them to adapt but no individual characteristics?

    Are you sure you thought this through?

  36. Corneel:
    Here is a tree of lifethat better represents modern understanding of relative diversity within Bacteria, Archaea and Eukarya. For giggles, try to locate “ancestors(a) to (g)” from your figure. Hopefully, you’ll realize that a fair amount of navelgazing went into its production.

    Both the tree I posted and the one you linked to are useful for understanding evolution. And both emphasise different aspects of diversity. Like the figure below, reality can be distorted as an aid to understanding and both trees should be interpreted in this way.

    The tree you linked to emphasises genomic diversity at the present time while downplaying the change over time that each branch undergoes. Although the length of the radiating to which we belong indicates that our lineage has had the greatest amount of evolutionary change over time. “Longer branches experienced more change”

    The tree I posted gives a clear indication of when the various groups first appeared in evolutionary time. The appearance of a group does not mean that it had no predecessors, it just means that it has undergone a dramatic change in morphology over the course of evolution.

    Both trees provide us with a lot of useful information.

    Corneel

    ETA: Also note how, in your figure, there is an implied progression from left to right. Phylogenetic trees can be rotated around their internal nodes without changing the topology, so why was it represented this way? Can you guess?

    The progression is not implied it is actual. Unless you believe that the advent of multicellularity is not a progression towards more organized complexity of individual organisms which leads to more autonomy from their milieu. For example endothermy allows for greater freedom of activity in environments where the temperature is constantly changing.

  37. CharlieM: Although the length of the radiating to which we belong indicates that our lineage has had the greatest amount of evolutionary change over time.

    How did you conclude that from an unrooted tree? Where do the main branches connect to the common ancestor, you reckon?

    CharlieM: The progression is not implied it is actual. Unless you believe that the advent of multicellularity is not a progression towards more organized complexity of individual organisms which leads to more autonomy from their milieu.

    Again, this is not something that follows from the figure you posted. For example, let’s consider node c (I believe this represents the branching off of bilaterians). Now, we are allowed to rotate the entire structure around this internal node like a crib mobile so let’s do that. Observe: humans are now cosying up side by side with corals and flatworms are top of the heap. Note that this tree is completely equivalent to the one that is depicted. So how did you conclude that there is a progression going from bacteria to humans based on that figure?

    You see what you want to see, Charlie.

  38. CharlieM,

    In the case of the peppered moth we can imagine two extreme forms diverging and remaining separate. But where will this lead? It is believed that moths existed on earth around 200 million years ago, so it is a fair assumption that these diverging peppered moths are descendants of lines of moths stretching back at least 200 million years. There is no evidence that, should they continue to exist, they will be anything but moths in the future. 

    ‘Moth’ is just what we call them. I’m willing to bet that in 200 million years both us and the word ‘moth’ will either have disappeared or changed beyond recognition. You play the same trick with ‘eukaryote’ in the subsequent comment to Corneel – imposing a static category simply by virtue of our nomenclature.

    We humans like to name stuff. Things are sufficiently static over our lifetimes that the names appear to attach to something persistent. But they don’t. We could go back to dinosaur times and declare ‘there is no evidence they will be anything but dinosaurs in the future’. But that sodding blackbird nicking my raspberries illustrates the dubiousness of that declaration.

  39. Allan Miller: ‘Moth’ is just what we call them.

    Yes, this another round of the naming game. No need to quantify stuff: As long as it bears the same name, it hasn’t really changed.

  40. Corneel: Yes, this another round of the naming game. No need to quantify stuff: As long as it bears the same name, it hasn’t really changed.

    Yep. Something I should have said – it’s only just occurred to me – is that going back in time, we’d presumably reach the common ancestor of butterflies and moths. It would probably have some flavour of both, as well as some distinctions of its own. It’s a … um … mutterfloth. A Charlie of that period would confidently declare that no amount of divergence from an unassuming population of mutterfloths would ever be anything but mutterfloths. Yet, the very fact of bifurcation has demanded two categories instead of one.

    If the butterfly lineage had failed to – ahem – take off, we’d just have modified mutterfloths, which we’d probably call primitive moths. But that’s not what happened. There are no mutterfloths any more. Except that there kind of are.

  41. Allan Miller: mutterfloth

    Haha. Do you want to hear something funny? Moths without butterflies is a paraphyletic group. Strict cladistically speaking, butterflies are a type of moth (Lepidoptera actually).

    They all look the same to Charlie, I bet. Mutterfloths, the lot of ’em.

  42. Allan Miller:
    CharlieM: Were the melanic forms not fixed in the population before selection due to tree discolouration caused by industrial pollution? Fluctuation in relative numbers of these previously fixed traits comes about due to selective pressures does it not?

    Allan Miller: The environmental change caused a change in the direction of selection. This does not mean that fluctuation is a component of selection, simply that selection is responsive to it.

    You talk about selection as if it were some overarching agent tweaking populations. Back in 2002 Craig Holdrege wrote about peppered moths and he had this to say:

    The problem is that the concept of selection includes at its core a selective agent. You can’t speak about selection without there being something doing the selecting. The “force” of natural selection is nothing more and nothing less than the specific agent(s) of selection; to speak of the force of natural selection in general terms is vacuous (and borders on the mystical). Until scientists know that predation by birds, the effects of air pollution on larvae, or other factors are at work, the concept of natural selection has no concrete content. Why, then, do both reviewers stress the fact of natural selection, when no agent of selection has been clearly determined?

    It’s as if at all costs they want to save natural selection and feel that if they lose it, they lose all grip on evolution. Natural selection is felt to be the best weapon against the threat of creationism, and if they don’t have this weapon, then what? But evolution — as opposed to a narrow, Darwinian interpretation of it — doesn’t stand or fall with the idea of natural selection; the evidence that organisms change over time is not at risk. Both Darwinians and creationists are stuck in ideologies — fixed mental frameworks through which they interpret the phenomena. Wouldn’t it be much more in the spirit of scientific inquiry to leave different avenues of explanation open as one continues to investigate the phenomena? Scientists should enjoy uncertainty and discovery as much as they enjoy fitting facts into a neat conceptual framework.

    If we are looking for the agents that have the greatest effect on relative percentages of the three moth varieties I would say predatory birds fit the bill. 🙂

    CharlieM: Say natural selection brought about the elimination of all peppered moths except the melanic forms. Surely this is a restriction of variability which will tend to make the species less capable of adapting to future changing conditions?

    Allan Miller: Sure. It happens. But it isn’t a universal. There is a constant flux of new mutations at one end, and a constant flux of fixation due to drift and selection at the other. So no biggie.

    But not all changes are equal in how easily they can be achieved. Fluctuating finch beaks and the plasticity of peppered moth populations are obvious observations. But the evolution of the simple proto feather to the Integumentary system of birds is a wondrous development that we can all see for ourselves if we are willing to learn as much as is possible by simply examining the feathers of birds.

    Believing in the creative power of mutations, drift and natural selection takes a great deal of faith.

  43. Allan Miller: Isn’t it fortunate that variation is, indeed, the rule? Except where it isn’t. We get along just fine with everyone having a nose, for example.

    Noses are wonderful things. The animal kingdom has examples of all sorts of extreme forms from the trunks of elephants to the blowholes of cetaceans. Human noses come in a variety of shapes and sizes and it mystifies me why some people undergo plastic surgery in order to possess a nose which conforms to some perceived idea of beauty. All they are doing is sacrificing something which displays their individual personality in order to display the most popular prominence of the times. And Micael Jackson was the proof that it doesn’t always succeed no matter how much money is thrown at it.

    So you have emphasized the point. Even if there is much variety in noses we can recognize a nose when we see one.

    Allan Miller: But here’s the thing: what would you call the response to changing environments, given population variation with the capacity to so respond? Huh? It’s got a name, you know…

    I would call it adaptive evolution. And I would class it as microevolution.

  44. Allan Miller: to Corneel,

    There is also a creative circularity in deprecating Natural Selection for its reduction of the variation whose principal purpose is to provide a response to … Natural Selection

    How can Natural Selection be something that can be deprecated? That would be like saying I disapprove of the process whereby stem cells give rise to specialist cells.

    I hereby disapprove of the oil filter in my car’s engine because it is selective in what it let’s past. We should campaign against particulate prejudice wherever we find it 🙂

  45. Alan Fox:
    CharlieM: In the case of the peppered moth we can imagine two extreme forms diverging and remaining separate. But where will this lead?

    Alan Fox: Speciation is the word you’re looking for

    I’m not looking for one-word answers. I think we’re all aware of the ambiguity of the species concept

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